Xingu–Tocantins–Araguaia Moist Forests
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The Xingu–Tocantins–Araguaia moist forests (NT0180) is an
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of ...
in the eastern Amazon basin. It is part of the Amazon biome. The ecoregion is one of the most severely degraded of the Amazon region, suffering from large-scale
deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated ...
and selective extraction of timber, particularly along the
Trans-Amazonian Highway The Trans-Amazonian Highway (official designation BR-230, official name Rodovia Transamazônica), was introduced on September 27, 1972. It is 4,000 km long, making it the third longest highway in Brazil. It runs through the Amazon forest and ...
and in the higher and more populated southern portions.


Location

The Xingu–Tocantins–Araguaia moist forests ecoregion has an area of . It lies to the south of the Amazon River in eastern Brazil. The
Xingu River The Xingu River ( ; pt, Rio Xingu, ; Mẽbêngôkre: ''Byti'', ) is a river in north Brazil. It is a southeast tributary of the Amazon River and one of the largest clearwater rivers in the Amazon basin, accounting for about 5% of its water. ...
defines the western boundary and the
Tocantins River The Tocantins River ( pt, Rio Tocantins, link=no , , Parkatêjê: ''Pyti'' ɨˈti is a river in Brazil, the central fluvial artery of the country. In the Tupi language, its name means " toucan's beak" (''Tukã'' for "toucan" and ''Ti'' for "beak ...
defines the eastern boundary. To the south the ecoregion rises into the Carajás Mountains, where it becomes interspersed with areas of seasonal forest and cerrado. Population centers include
São Félix do Xingu São Félix do Xingu is a municipality in the state of Pará in the Northern region of Brazil. The city is served by São Félix do Xingu Airport. With an area of , it is the third largest municipality in Pará and the sixth large ...
on the Xingu, Porto de Moz,
Oeiras do Pará Oeiras do Pará is a municipality in the state of Pará in the Northern region of Brazil. See also *List of municipalities in Pará This is a list of the municipalities in the state of Pará (PA), located in the North Region of Brazil. Pará ...
and Gurupá on the Amazon, and Marabá on the Tocantins. The
Marajó várzea The Marajó várzea (NT0138) is an ecoregion of seasonally and tidally flooded várzea forest in the Amazon biome. It covers a region of sedimentary islands and floodplains at the mouth of the Amazon that is flooded twice daily as the ocean tides ...
at the mouth of the Amazon lies to the north. To the west, on the other side of the Xingu River, is the
Tapajós–Xingu moist forests The Tapajós–Xingu moist forests (NT0168) is an ecoregion in the eastern Amazon basin. It is part of the Amazon biome. The ecoregion extends southwest from the Amazon River between its large Tapajós and Xingu tributaries. Location The Tapajó ...
. To the northeast is the
Tocantins–Araguaia–Maranhão moist forests The Tocantins–Araguaia–Maranhão moist forests (NT0170), also called the Tocantins/Pindaré moist forests, is an ecoregion in the north of Brazil to the south of the mouth of the Amazon River. It is part of the Amazon biome. The ecoregion con ...
. To the southeast and south the ecoregion blends into the Mato Grosso seasonal forests ecoregion, and in some places directly adjoins the
Cerrado The ''Cerrado'' (, ) is a vast ecoregion of tropical savanna in eastern Brazil, particularly in the states of Goiás, Mato Grosso do Sul, Mato Grosso, Tocantins, Minas Gerais, and the Federal District. The core areas of the Cerrado biome are the ...
ecoregion.


Physical

The terrain is undulating. In many areas the soils are rich, but in some parts they are low in nutrients. The north of the ecoregion is on the lowland Amazon basin, while in the south it rises into the higher ground of the Brazilian Shield. Elevations range from sea level at the Amazon to in the Carajás Mountains in the south. The forest contains a network of
blackwater river A blackwater river is a type of river with a slow-moving channel flowing through forested swamps or wetlands. As vegetation decays, tannins leach into the water, making a transparent, acidic water that is darkly stained, resembling black tea ...
s, stained dark with
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'' ...
s and holding little suspended sediment. It contains the middle and lower basin of the Araguaia River, a tributary of the Tocantins. There are many smaller blackwater rivers including the Pacajá and Anapu tributaries of the Amazon River, the Bacajá and Fresco tributaries of the Xingu, and the
Parauapebas Parauapebas is a municipality in the state of Pará in the Northern region of Brazil. Geography To the north is the municipality of Marabá; to the east, Curionópolis; to the south-east, Canaã dos Carajás; to the south, Água Azu ...
and Catete tributaries of the Tocantins.


Ecology

The ecoregion is in the
Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeo ...
and the
tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discon ...
biome.


Climate

The
Köppen climate classification The Köppen climate classification is one of the most widely used climate classification systems. It was first published by German-Russian climatologist Wladimir Köppen (1846–1940) in 1884, with several later modifications by Köppen, notabl ...
is "Am": equatorial, monsoonal. Temperatures are fairly steady throughout the year, slightly higher in April and slightly lower in July. Average temperatures range from , with a mean of . The region is drier than the ecoregions further to the west. In the middle Xingu annual rainfall is . Elsewhere average annual precipitation is about Monthly precipitation ranges from in July to in March.


Flora

The diverse terrain results in diverse flora and fauna, particularly on the richer soils, with many endemic species. Most of the original vegetation is evergreen tropical rainforest on terra firme. In the north there is lowland Amazon forest up to high, graduating to dense submontane forest and then open submontane forest further south. There are fewer epiphytes and
orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowerin ...
s in these forests than in the ecoregions further west. There are areas of liana forests on higher ground in the south and southeast and of forests dominated by babaçu palm ( Attalea genus) and/or
Brazil nut The Brazil nut (''Bertholletia excelsa'') is a South American tree in the family Lecythidaceae, and it is also the name of the tree's commercially harvested edible seeds. It is one of the largest and longest-lived trees in the Amazon rainforest. ...
(''Bertholletia excelsa''). The stands of babaçu palms are thought to be originally planted by humans. The blackwater river banks are lined with white-sand
igapó Igapó (, from Old Tupi: "root forest") is a word used in Brazil for blackwater-flooded forests in the Amazon biome. These forests and similar swamp forests are seasonally inundated with freshwater. They typically occur along the lower reaches of ...
flooded forests. In the south the ecoregion is broken up by areas of savanna woodlands from the
cerrado The ''Cerrado'' (, ) is a vast ecoregion of tropical savanna in eastern Brazil, particularly in the states of Goiás, Mato Grosso do Sul, Mato Grosso, Tocantins, Minas Gerais, and the Federal District. The core areas of the Cerrado biome are the ...
biome of the Central Brazilian Plateau. The liana forests cover large areas of comparatively rich soil. They are more open than the humid terra firme forest, with a lower canopy of or less. They contain many huge lianas at all levels from the families
Bignoniaceae Bignoniaceae is a family of flowering plants in the order Lamiales commonly known as the bignonias or trumpetvines.Vernon H. Heywood, Richard K. Brummitt, Ole Seberg, and Alastair Culham. ''Flowering Plant Families of the World''. Firefly Book ...
, Fabaceae, Hippocrateaceae,
Menispermaceae Menispermaceae (botanical Latin: 'moonseed family' from Greek ''mene'' 'crescent moon' and ''sperma'' 'seed') is a family of flowering plants. The alkaloid tubocurarine, a neuromuscular blocker and the active ingredient in the 'tube curare' form ...
, Sapindaceae, and Malpighiaceae. They also hold large trees including '' Apuleia molaris'', '' Bagassa guianensis'', '' Caryocar villosum'', '' Hymenaea parvifolia'', '' Tetragastris altissima'', ''
Astronium graveolens ''Astronium graveolens'' is a species of flowering tree in the cashew family, Anacardiaceae, that is native to the Yucatán Peninsula in Mexico, Central America, and South America as far south as Bolivia. Common names include glassywood, ronrón ...
'', ''
Astronium lecointei ''Astronium lecointei'' (Portuguese common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the nor ...
'', '' Apuleia leiocarpa'' var. molaris, '' Sapium marmieri'', '' Acacia polyphylla'' and '' Elizabetha''. There are many endemic species such as '' Cenostigma tocantinum'', '' Ziziphus itacaiunensis'' and '' Bauhinia bombaciflora''. Typical trees in the middle Tocantins upland include '' Cenostigma tocantinum'', '' Bombax tocantimumi'', '' Bombax macrocalyx'', '' Matisia bicolor'', '' Strychnos melinoniana'' and '' Strychnos solimoesana''. '' Swietenia macrophylla'' is found in the moist areas.


Fauna

There are 153 species of mammals, a relatively low number for the Amazon region. Of these 90 are bats and 21 are rodents. There are 8 species of primates including the
white-nosed saki The white-nosed saki (''Chiropotes albinasus'') is a species of bearded saki, a type of New World monkey, endemic to the south-central Amazon rainforest in Brazil. Both its scientific and common name were caused by the authors working from dead ...
(''Chiropotes albinasus''),
red-handed tamarin The golden-handed tamarin (''Saguinus midas''), also known as the red-handed tamarin or Midas tamarin, is a New World monkey belonging to the family Callitrichidae. Distribution and habitat This species is native to wooded areas north of the Ama ...
(''Saguinus midas''),
red-bellied titi The red-bellied titi monkey or dusky titi (''Plecturocebus moloch'') is a species of titi monkey, a type of New World monkey, endemic to Brazil. It lives in forests and thickets. It has a rounded head and a thick, soft coat and frequently adopts ...
(''Callicebus moloch''), Azara's night monkey (''Aotus azarae infulatus'') and white-cheeked spider monkey (''Ateles marginatus''). The last two are endemic to this ecoregion and the Tapajós-Xingu moist forests to the west. Other species of mammals include
white-lipped peccary The white-lipped peccary (''Tayassu pecari'') is a species of peccary found in Central and South America and the only member of the genus ''Tayassu''. Multiple subspecies have been identified. White-lipped peccaries are similar in appearance to ...
(''Tayassu pecari''),
collared peccary The collared peccary (''Dicotyles tajacu'') is a species of artiodactyl (even-toed) mammal in the family Tayassuidae found in North, Central, and South America. It is the only member of the genus ''Dicotyles''. They are commonly referred to as ...
(''Pecari tajacu''), cougar (''Puma concolor''), jaguar (''Panthera onca''),
South American tapir The South American tapir (''Tapirus terrestris''), also commonly called the Brazilian tapir (from the Tupi ''tapi'ira''), the Amazonian tapir, the maned tapir, the lowland tapir, the ''anta'' (Portuguese), and ''la sachavaca'' (literally "bushco ...
(''Tapirus terrestris''), brocket deer (genus ''Mazama''),
bare-tailed woolly opossum The bare-tailed woolly opossum (''Caluromys philander'') is an opossum from South America. It was first described by Swedish zoologist Carl Linnaeus in 1758. The bare-tailed woolly opossum is characterized by a gray head, brown to gray coat, or ...
(''Caluromys philander''), and
seven-banded armadillo The seven-banded armadillo (''Dasypus septemcinctus''), also known as the Brazilian lesser long-nosed armadillo, is a species of armadillo from South America found in Paraguay, Argentina, Bolivia and Brazil. It is a solitary nocturnal, terrest ...
(''Dasypus septemcinctus''). Endangered mammals include white-cheeked spider monkey (''Ateles marginatus''), black bearded saki (''Chiropotes satanas'') and
giant otter The giant otter or giant river otter (''Pteronura brasiliensis'') is a South American carnivorous mammal. It is the longest member of the weasel family, Mustelidae, a globally successful group of predators, reaching up to . Atypical of muste ...
(''Pteronura brasiliensis''). 527 species of birds have been recorded including many species of
egret Egrets ( ) are herons, generally long-legged wading birds, that have white or buff plumage, developing fine plumes (usually milky white) during the breeding season. Egrets are not a biologically distinct group from herons and have the same build ...
, heron, hawk and
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
along the rivers and meadows. Other birds include Brigida's woodcreeper (''Hylexetastes perrotii brigidai''),
hyacinth macaw The hyacinth macaw (''Anodorhynchus hyacinthinus''), or hyacinthine macaw, is a parrot native to central and eastern South America. With a length (from the top of its head to the tip of its long pointed tail) of about one meter it is longer tha ...
(''Anodorhynchus hyacinthinus''),
scarlet macaw The scarlet macaw (''Ara macao'') is a large red, yellow, and blue Central and South American parrot, a member of a large group of Neotropical parrots called macaws. It is native to humid evergreen forests of the Neotropics. Its range extends from ...
(''Ara macao''), many parrots (genera ''
Amazona Amazon parrots are parrots in the genus ''Amazona''. They are medium-sized, short-tailed parrots native to the Americas, with their range extending from South America to Mexico and the Caribbean. ''Amazona'' is one of the 92 genera of parrots t ...
'' and ''
Pionus ''Pionus'' is a genus of medium-sized parrots native to Mexico, and Central and South America. Characteristic of the genus are the chunky body, bare eye ring (which can vary in color), and short square tail. They are superficially similar to Ama ...
'') and parakeets (genera ''
Aratinga ''Aratinga'' is a genus of South American conures. Most are predominantly green, although a few are predominantly yellow or orange. They are social and commonly seen in groups in the wild. In Brazil, the popular name of several species usually is ...
'', ''
Pyrrhura ''Pyrrhura'' (Greek Red/Fire Tail) is a genus of parrots in the Arini tribe. They occur in tropical and subtropical South America and southern Central America (Panama and Costa Rica). Most are restricted to humid forest and adjacent habitats, ...
'' and ''
Brotogeris ''Brotogeris'' is a genus of small parrots endemic to Central and South America. Their closest relatives are the monk parakeet and the and the cliff parakeet in the genus '' Myiopsitta''. They eat seeds and fruit. The word ''brotogeris'' means " ...
). Endangered birds include green-thighed parrot (''Pionites leucogaster''), red-necked aracari (''Pteroglossus bitorquatus'') and yellow-bellied seedeater (''Sporophila nigricollis''). The rivers are home to spectacled caiman (''Caiman crocodilus''), black caiman (''Melanosuchus niger''),
yellow-spotted river turtle The yellow-spotted Amazon river turtle (''Podocnemis unifilis''), also known commonly as the yellow-headed sideneck turtle and the yellow-spotted river turtle, and locally as the taricaya, is one of the largest South American river turtles. It ...
(''Podocnemis unifilis''), Amazonian manatee (''Trichechus inunguis''),
Amazon river dolphin The Amazon river dolphin (''Inia geoffrensis''), also known as the boto, bufeo or pink river dolphin, is a species of toothed whale classified in the family Iniidae. Three subspecies are currently recognized: ''I. g. geoffrensis'' (Amazon river ...
(''Inia geoffrensis'') and
tucuxi The tucuxi (''Sotalia fluviatilis''), alternatively known in Peru ''bufeo gris'' or ''bufeo negro'', is a species of freshwater dolphin found in the rivers of the Amazon basin. The word ''tucuxi'' is derived from the Tupi language word ''tuchu ...
(''Sotalia fluviatilis'').


Status

The
World Wildlife Fund The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wo ...
classes the ecoregion "Vulnerable". The Tapajós-Xingu, Xingu-Tocantins-Araguaia and Tocantins/Pindare moist forests ecoregions on the eastern edge of the Amazon basin have all been badly affected by human settlement and deforestation. The Xingu-Tocantins-Araguaia region is the most deforested and degraded part of the Amazon region after the adjoining Tocantins/Pindare moist forests to the east. It is crossed by the
Trans-Amazonian Highway The Trans-Amazonian Highway (official designation BR-230, official name Rodovia Transamazônica), was introduced on September 27, 1972. It is 4,000 km long, making it the third longest highway in Brazil. It runs through the Amazon forest and ...
(BR-230) from east to west, and by the BR-422 highway along the Tocantins. Most of the land along these roads has been deforested and replaced by pasture or farm fields. The remaining forest has been stripped of its valuable trees, and many native fauna and flora have become locally extinct. Large-scale mining near Marabá and elsewhere have directly affected the environment, and demand large quantities of wood to run the smelters. During the period from 2004 to 2011 the ecoregion experienced an annual rate of habitat loss of 0.94%. Global warming will force tropical species to migrate uphill to find areas with suitable temperature and rainfall. Low, flat, deforested ecoregions such as the Xingu-Tocantins-Araguaia moist forests are extremely vulnerable. Conservation units include the Tapirapé Biological Reserve and
Tapirapé-Aquiri National Forest Tapirapé-Aquiri National Forest ( pt, Floresta Nacional do Tapirapé-Aquiri) is a national forest in the state of Pará, Brazil. It is surrounded by other protected areas, so has suffered relatively little from deforestation. Location The Tapir ...
.


Notes


Sources

* * * * * * {{DEFAULTSORT:Xingu-Tocantins-Araguaia moist forests Ecoregions of Brazil Amazon biome Neotropical tropical and subtropical moist broadleaf forests